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12–––3––––––43OOOOHOHHHOHHOHO53OOOOHOHHHOHHOHO63OOOOHOHHHOHHOHO7–891011–12––13–1414β-OH14α-OHD3β-OH3α-OH17αβ-α−3β-OHβ-1,4OHHOHHOHOO15–3OOOOHOHHHOHHOHO16