FoPSIIFmPSIIFoFo=1/(1/Fo-1/Fm+1/FmFmPSIIFFv/FmandY(II)PSIIFv/FmY(II)PSIIFv/FmPSIIY(II)Y(II)Y(II)qPandqLPSIIqPpuddlemodelSchreiberetal.1986asformulatedbyvanKootenandSnel,1990qLlakemodel,Krameretal.2004qNandNPQY(NO)andY(NPQ)Kramer2004Y(NPQ)PSIIY(NPQ)YNPQY(NO)PSIIY(NO)YNOY(II)+Y(NO)+Y(NPQ)=1