1EvolutionaryGames13Chapter13Slide2EvolutionaryGamesWehavesofarstudiedgameswithmanydifferentfeatures:SimultaneousandsequentialmovesZero-sumandnon-zero-sumpayoffsStrategicmovestomanipulaterulesofgamestocomeOne-shotandrepeatedplayGamesofcollectiveactioninwhichalargenumberofpeopleplaysimultaneously2Slide3EvolutionaryGames……Alltheplayersinallthesegamesarerational:eachplayer…………hasaninternallyconsistentvaluesystem……cancalculatetheconsequencesofherstrategicchoices……makeschoicethatbestfavorsherinterestsSlide4EvolutionaryGames……Onepossiblealternativetorationalitycanbefoundinthebiologicaltheoryofevolutionandevolutionarydynamics,where…………goodstrategieswillberewardedwithhigherpayoffs……playerscanobserveorimitatesuccessandexperimentwithnewstrategies……goodstrategieswillbeusedmoreoftenandbadstrategieslessoften,asplayersgainexperienceplayingthegame.3Slide5Outline*Theframework**Prisoners’dilemmaChickenTheassurancegame*InteractionsacrossspeciesThehawk-dovegame*ThreePhenotypesinthePopulationSomeGeneralTheoryPlayingthefieldSlide6TheFrameworkTheprocessofevolutioninbiologyoffersaparalleltothetheoryofgamesusedbysocialscientists.Thistheoryrestsonthreefundamentals:HeterogeneityFitnessSelection4Slide7TheFrameworkAsignificantpartofanimalbehaviorisgenericallydetermined;acomplexofoneormoregenes(genotype)governsaparticularpatternofbehavior,calledabehaviorphenotype.ExamplesAerodynamiccharacteristicsofabird’swingsAggressiveorcooperativebehaviorLocationsofnestingsitesSlide8TheFrameworkNaturaldiversityofthegenepoolensuresaheterogeneityofphenotypesinthepopulation.Somebehaviorsarebettersuitedthanotherstotheprevailingconditions,andthesuccessofaphenotypeisgivenaquantitativemeasurecalleditsfitness.5Slide9TheFrameworkThefitterphenotypesthenbecomerelativelymorenumerousinthenextgenerationthanthelessfitphenotypes.Thisprocessofselectionisthedynamicthatchangesthemixofgenotypesandphenotypesandperhapsleadseventuallytoastablestate.Slide10TheFrameworkFromtimetotime,chanceproducesnewgeneticmutations.Manyofthesemutationsproducebehavior(thatis,phenotypes)thatareillsuitedtotheenvironment,andtheydieout.Butoccasionallyamutationleadstoanewphenotypethatisfitter.Thensuchamutantgenecansuccessfullyinvadeapopulation–thatis,spreadtobecomeasignificantproportionofthepopulation.6Slide11TheFrameworkBiologistscallaconfigurationofapopulationanditscurrentphenotypesevolutionarystableifthepopulationcannotbeinvadedsuccessfullybyanymutant.Thisisastatictest;butoftenamoredynamiccriterionisapplied:aconfigurationisevolutionarystableifitisthelimitingoutcomeofthedynamicofselection,startingfromanyarbitrarymixtureofphenotypesinthepopulation.Slide12TheFrameworkThefitnessofaphenotypedependsontherelationshipoftheindividualorganismtoitsenvironment.Italsodependsonthewholecomplexoftheproportionsofdifferentphenotypesthatexistintheenvironment.Forourpurpose,thisinteractionbetweenphenotypeswithinaspeciesisthemostinterestingpartofthestory.7Slide13TheFrameworkThebiologicalprocessofevolutionfindsareadyparallelingametheory.Phenotypevs.StrategyFitnessvs.PayoffsSlide14TheFrameworkBecausethepopulationisamixofphenotypes,differentpairsselectedfromitwillbringtotheirinteractionsdifferentcombinationsofstrategies.Theactualquantitativemeasureofthefitnessofaphenotypeistheaveragepayoffthatitgetsinallitsinteractionswithothersinthepopulation.8Slide15TheFrameworkThetheoryofevolutionarygamesseemsaready-madeframeworkforanewapproachtogametheory,relaxingtheassumptionofrationalbehavior.Theideaofinheritanceofstrategiescanbeinterpretedmorebroadlyinapplicationsofthetheoryotherthanbiology.Slide16TheFramework“”Thereasonsthatthefitterstrategiesproliferateandthelessfitonesdieoutinsocioeconomicgamesdiffersfromthestrictgeneticmechanismofbiology:ObservationsandImitationsPurposivethinkingandrevisionofpreviousrulesofthumbConsciousexperimentation9Slide17TheFrameworkWhyaplayerplayssuchastrategy?RationalchoicesGeneticsSocialization,culturalbackground,educationsAruleofthumbbasedonpastexperienceWillsocietyendupwithasituationinwhichallpoliticiansareconcernedwithreelection,andallfirmswithprofit?Slide18TheFrameworkEvolutionarystableconfigurationsofbiologicalgamescanbeoftwokinds.Monomorphism:Asinglephenotypeprovesfitterthananyothersandthepopulationcomestoconsistofitalone.Inthiscase,theuniqueprevailingstrategyiscalledanevolutionarystablestrategy(ESS).Polymorphism:Twoormorephenotypesareequallyfit(andfitterthansomeothersnotplayed);sotheymaybeabletocoexistincertainproportions.10Slide19TheFrameworkThewholeset-upwhichconstitutesanevolutionarygameis:ThePopulationItsconceivablecollectionofphenotypesThepayoffsmatrixintheinteractionsofthephenotypesTherulefortheevolutionofpopulationproportionsofthephenotypesinrelationtotheirfitnessAnevolutionarystableconfigurationofthepopulationcanbecalledanequilibriumoftheevolutionarygame.Slide20Prisoners’DilemmaSupposeapopulationismadeupoftwophenotypes:cooperators,defectors.Eachindividual(eitheracooperatororadefectors)inthepopulationischosenatrandomtocompeteagainstanotherrandomrival.11S